Your mishap has rippled into the Duffer household in that my WM280 Manual has disappeared. Only a small thing in comparison but my blood pressure has gone through the roof.
I was going to look at the exploded parts diagram to see if I could understand what stops the self-act engaging when the half-nut is closed. On my machine it feels like a deliberate interlock. I wonder if moving your lathe into the cellar, perhaps tilting it, caused the interlock to engage and that dropping the apron to look inside freed it.
If it's any consolation at all, many thanks for sharing what you've done about the problem. Now I've seen the photos of your lathe in bits I'm much more confident about servicing mine should the need arise!
Hope you have it working soon and that the problem never recurs.
Dave